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Carabidae
Elytron lateral margin
unmodified
Plica present
Maxillary Palpomere 4
Subulate
Obliquely attached to palpomere 3
unmodified
Triangular
Inner hind tibial spur
Subequal to outer spur
Long and serrulate
Transverse row of setae on vertex
Absent
Present
Mesocoxal closure
Ventrites disjunct
Ventrites conjunct
Antennal Insertion Location
In line with mandibular bases
On frons, between mandibular bases
Sutural interval
Not flatened, normal
Flattened, distinctly pinched
Pronotum lobed basally
Present
Absent
Vestiture of setae
Present
Absent
Setigerous puncture in scrobe
Setigerous puncture present
Setigerous puncture absent
Species Count: 663
Carabidae
Rhysodidae
Trachypachidae