Scales of stems and petiole bases mostly bicolored with broad, usually continuous, dark central stripe, scales narrowly lanceolate. Pinnae with longest hairs composed of 5--8 cells. Indusial segments broad, not at all filamentous. Spores averaging 39--46 µm. 2 n = 76. Sporulating summer--fall. Shaded cracks and ledges on cliffs; mostly subacidic shale and sandstone; 300--1700 m; Ark., Ky., N.C., Tenn., Va., W.Va.